Traditional Culture Encyclopedia - Weather inquiry - Complete details of mudskippers.
Complete details of mudskippers.
Mud-coated fish are the genius of fish. They spend a long time in the water. The place where they live is full of mangroves, and they are happy to climb the trunk or branches. They use ventral fins as suckers to catch trees and pectoral fins to climb up. There are small mouths around the gills of mudskippers, which can hold water for one breath, much like we can hold water for one breath. They climb trees, so they can stay out of the water at high tide.
Basic introduction Chinese name: mudskipper Latin scientific name: Periophthalmus? Modesto? Nicknames: Stripe, Jumping Fish, Periodontitis. Two names: Periodontitis? Modestus domain: Animal kingdom: Chordata subfamily: Vertebrate subfamily: Actinidia subfamily: Perciformes: gobies subfamily: mudskippers subfamily: mudskippers subfamily: mudskippers subfamily: mudskippers subfamily: mudskippers distribution area: mangrove wetlands common in Hongkong, Taiwan Province Province and Southeast Asia. English name: basic information, morphological characteristics, living environment, living habits, reproduction, underground caves, courtship dance, spawning room, towers and trenches, species classification, geographical distribution, artificial breeding, pest control, fishing, main value, related news, basic information, common name of mudskipper, alias jumping fish. Bony fish of perciformes Small fish of warm waters and temperate coastal areas. Widely distributed in the tropical and subtropical inshore shallow waters of the west coast of Africa, Indo-Pacific waters, Australia and New hebrides, but not in America. It is distributed along the coast of China. Morphological characteristics The body of mudskipper is cylindrical, with a general body length of10 ~ 20cm and a weight of 20 ~ 50g. The eyes are small, protruding above the rear edge of the head, the lower eyelids are developed, and the mouth is in the terminal position. Each jaw has/kloc-0 rows of teeth, the maxillary teeth are tapered, and the three teeth on each side in front are canine teeth. The mandibular teeth are tilted outward and almost lying down. The back is dark brown. The abdomen is gray. There are 6 black stripes on the back, irregular green-brown spots on the whole body and 2 dorsal fins. 1 The dorsal fin is very small, with only five fin spines, and the ends of the fin spines are filiform, of which the third spine is the longest; The second dorsal fin and gluteal fin are very long, with three long gray-white horizontal lines, which are roughly equal in length. The caudal fin is wedge-shaped and wide; The pectoral fin has a yellow-green insect pattern. Mature individuals is generally 80 ~ 90 mm long and weighs 15 ~ 25g. Adults can reach 120 ~ 150 mm in length and weigh 30 ~ 40 grams. The side of the body is flat, the trailing edge is straight, the abdominal edge is shallow arc, the tail handle is high, and the body and head are covered with small round scales. The first dorsal fin is quite high, the base is short, and the five-fin spine is slender and filiform; The fin of the second dorsal fin is lower, reaching the base of the caudal fin, the ventral fin is short, and the pectoral fin and caudal fin are pointed and round. The body is blue-brown or gray-brown, and there are 6 ~ 7 gray-black horizontal stripes along the base of dorsal fin on the upper side of the body. There are many bright blue dots scattered on the side and head, the abdomen is white, the first dorsal fin is dark blue, the second dorsal fin is blue-gray, the ventral fin is light yellow, and the caudal fin is gray-black. Male and female mudskippers are indistinguishable in appearance, but the female reproductive pore is large, round and red. Male reproductive pores are long and narrow. Living environment inhabits the salty and fresh water area of the estuary, the coastal beach or the low tide area of sediment deposition, and has strong tolerance to the water quality of the harsh environment. Euryhaline likes to live in caves, which are generally Y-shaped and consist of channels, anterior orifices and posterior orifices. The front hole is used for access and the rear hole is used for ventilation. Use pectoral fin and tail handle to crawl or jump on water, beach and rock; Crawling in the mud, covered with benthic diatoms, cyanobacteria, fish food, and a small amount of copepods and organic matter. The mudskipper can use pectoral fins and tail stalks to crawl and jump on the beach. The skin and tail can be used as auxiliary respiratory organs, and it can live out of the water for a long time, and its tolerance to harsh environment is stronger than that of ordinary fish. This kind of fish has the habit of drilling holes to inhabit, and its passage has at least two openings, one is the front opening and the other is the back opening, and the front opening is larger. The channel is irregular, generally Y-shaped, and its depth varies with soil quality, reaching 50 ~ 70 cm. Generally speaking, mudskippers live alone and can hatch in tunnels during the spring and summer breeding season. Mudskippers can use pectoral fins and caudal fins to crawl or jump on water, beaches and rocks. The inner gill cavity, skin and tail of mudskipper can be used as respiratory AIDS. As long as your body is wet, you can live out of water for a long time. It is a temperate, wide-temperate and euryhaline fish. Muddy fish will die if they stay in fresh water for more than ten days. When the water temperature is 14 ~ 30℃, it grows fast, and when it is below 14℃, it lives in seclusion in caves, requiring less food and growing slowly. Mud-coated fish have stronger tolerance to water quality in harsh environment than ordinary fish, and have the same habits as freshwater eels. They can climb into rivers and inhabit, and countless small holes can be seen scattered in the beach. Generally, there are more than two holes in the channel, one is the front hole and the other is the rear hole (commonly known as the rear bubble hole). The front mouth is the access artery, and the back mouth is the smooth water flow and air circulation. The holes in caves are irregular, usually in the shape of "Y" or "U". The depth and length of the channel vary according to the properties of the substrate. Soft clay caves are deep and long, reaching 50 ~ 70 cm. Generally speaking, holes are exclusive. But in the breeding season, hermaphroditism often occurs. The holes of mudskippers are also used as spawning rooms. Bullet fish live in coastal areas and tidal flats below estuaries. They jump out of holes on sunny days, feed on mudflats, and feed on benthic algae, insects and other small creatures on mudflats. According to the analysis of stomach contents, mudskippers mainly eat attached diatoms. At low tide or at the bottom of the drained pond, it is often seen that mudskippers feed on benthic algae, that is, their jaws touch the beach surface, and their heads swing left and right like plowing, crawling forward and feeding on benthic algae. Mud-wrapped fish have no bad habit of killing each other. Artificial culture uses fertilization to cultivate basic bait and promote the growth of loach. After low tide, mudskippers often face the danger of being preyed by birds and various terrestrial mammals, and underground caves provide them with a safe environment. After the high tide, mudskippers can hide in their caves to avoid the attacks of various carnivorous fish that come to the shallow beach to feed. The cave of the mudskipper can be used not only as a refuge, but also as a nursery. Caves are very important for the safety of mudskippers, but the caves of mudskippers are also facing danger-the water in caves is often seriously deprived of oxygen. In this regard, females and males will continue to swallow air in turn and inject it into their holes, so as to establish an underground airbag to alleviate hypoxia. Bullet fish have the habit of eating out of water. Whenever the tide ebbs, it often crawls on the mud for food, or climbs on rocks and mangroves to catch insects, or climbs on rocks to bask in the sun. When it comes out of the water, the developed gill cavity is filled with air, and the tail is immersed in water to assist breathing. Living without water has become an important habit of it. It can move as lively as a lizard on land, and the pectoral fin muscle stalk can move back and forth freely, playing the role of a reptile forelimb. In order to strengthen the ability of crawling on land, the gluteal fin becomes very low and the fin of the lower lobe of the caudal fin becomes thicker. When the pectoral fin moves forward, the pelvic fin plays a role in supporting the body. When it jumps in a short distance, it only depends on the activity of pectoral fins; When jumping more than one meter, you must beat the ground with your tail to assist. And it only jumps long distances when it is anxious or afraid. At low tide, mudskippers jump around and chase each other. Living habits Pinball fish have the habit of leaving water for food. Whenever the tide ebbs, it often crawls on the mud for food, or climbs on rocks and red bushes to catch insects, or climbs on rocks to bask in the sun. When it comes out of the water, the developed gill cavity is filled with air, and the tail is immersed in water to assist breathing. Living without water has become an important habit of it. It can move as lively as a lizard on land, and the pectoral fin muscle stalk can move back and forth freely, playing the role of a reptile forelimb. In order to strengthen the ability of crawling on land, the gluteal fin becomes very low and the fin of the lower lobe of the caudal fin becomes thicker. When the pectoral fin moves forward, the pelvic fin plays a role in supporting the body. When it jumps in a short distance, it only depends on the activity of pectoral fins; When jumping more than one meter, you must beat the ground with your tail to assist. And it only jumps long distances when it is anxious or afraid. At low tide, mudskippers jump around and chase each other. The breeding season of mudskippers is from April to September every year, and each parent fish can lay about 10000 eggs. Oval position is spherical, sticky and yellow. In summer and autumn, a large number of fry can be seen in the estuary area where the specific gravity of seawater is lower than 1.05438+05. Its growth rate is slow, and it generally takes 1 ~ 2 years from seedling to adult fish.
Mature individuals of mudskipper is12 ~15cm, and its weight is 30 ~ 40g. The breeding season is from April to September, and the peak period is from May to July. Lay eggs naturally in caves on the beach, ovulate in them, hide the fertilized eggs deeply and naturally hatch. During sexual maturity, the reproductive pores of female fish are red and swollen, large and round; The male fish's reproductive pores are narrow and elongated in a pointed shape. The ovary is yellow, and the eggs laid are thick, round, yellowish and transparent. The female fish has 65,438+0 ~ 25,000 eggs, and one end of the egg membrane has a cluster of attachment filaments, which can be attached to other objects. The egg diameter was 0.565,438+0 ~ 0.665,438+0 mm, and the fertilized egg was incubated for 87 hours at the temperature of 26.5 ~ 29.0℃ and the salinity of 25 ‰ ~ 27 ‰. There are two seminal vesicles in male fish, which are reddish and strip-shaped on the ventral side. The mudskipper grows rapidly, and the early larvae have good environmental conditions. The benthic algae enrichment area can grow to 12 ~ 13 cm in that year, reaching the commercial specifications. Generally, it can meet the commodity specifications after two years. The average life span of mudskippers is 3 ~ 5 years, and the longest life span is 7 years. We still don't know the important role of towers and trenches in biology. We suspect that trenches may be used to prevent other animals from approaching their caves, and the tower may be an observation point from which we can observe whether there are prey or carnivores outside the caves. In the morning, we found the mudskipper lazily lying in the tower-shaped hole. At this time, the air diffuses 7 degrees Celsius, which is higher than the temperature of the mudskipper hole. Mud-coated fish are preheated by the hole to prepare for the day's activities. The breeding season of mudskippers is from April to September every year, and each parent fish can lay about 65,438+00000 eggs. Oval position is spherical, sticky and yellow. In summer and autumn, a large number of fry can be seen in the estuary area where the specific gravity of seawater is lower than 1.05438+05. Its growth rate is slow, and it generally takes 1 ~ 2 years from seedling to adult fish. Mature individuals of mudskipper is12 ~15cm, and its weight is 30 ~ 40g. The breeding season is from April to September, and the peak period is from May to July. Lay eggs naturally in caves on the beach, ovulate in them, hide the fertilized eggs deeply and naturally hatch. During sexual maturity, the reproductive pores of female fish are red and swollen, large and round; The male fish's reproductive pores are narrow and elongated in a pointed shape. The ovary is yellow, and the eggs laid are thick, round, yellowish and transparent. The female fish has 65,438+0 ~ 25,000 eggs, and one end of the egg membrane has a cluster of attachment filaments, which can be attached to other objects. The egg diameter was 0.565,438+0 ~ 0.665,438+0 mm, and the fertilized egg was incubated for 87 hours at the temperature of 26.5 ~ 29.0℃ and the salinity of 25 ‰ ~ 27 ‰. There are two seminal vesicles in male fish, which are reddish and strip-shaped on the ventral side. The mudskipper grows rapidly, and the early larvae have good environmental conditions. The benthic algae enrichment area can grow to 12 ~ 13 cm in that year, reaching the commercial specifications. Generally, it can meet the commodity specifications after two years. The average life span of mudskippers is 3 ~ 5 years, and the longest life span is 7 years. Although this kind of fish that can move on land attracted the attention of some naturalists as early as 300 years ago, not many people can understand the underground life style of this fish until now. Although mudskippers can leave the water and wander alone on the flat shallow beach, for many mudskippers, an underground nest full of water may be more important for their survival. They can lurk in the cave, waiting for an opportunity to launch a sudden attack on the prey outside the cave. When they are threatened by predators, they can retreat quickly. After low tide, mudskippers often face the danger of being preyed by shorebirds and various terrestrial mammals, and underground caves provide them with a safe environment. After the high tide, mudskippers can hide in their caves to avoid the attacks of various carnivorous fish coming to shallow beaches. The cave of the mudskipper can be used not only as a refuge, but also as a nursery. Caves are very important for the safety of mudskippers, whether adults or larvae, but the caves of mudskippers are also facing danger-the water in caves is often seriously deprived of oxygen. Bullet fish must be able to tolerate or overcome the lack of oxygen in caves, not only adult fish, but also their developing eggs. Because both high tide and low tide will destroy the cave of mudflat fish, mudflat fish must pull out the sand and stones in the cave with their mouths whenever the tide is low. Courtship Dance Every spring, the male fish will find a suitable ground to divide his sphere of influence, and then dig a hole two feet (0.6 meters) deep in the mud. The hole is shaped like a "J" (sometimes it will dig a "Y"-shaped hole with two entrances), and the lower layer in the hole is its spawning room, which is used to store their eggs. After digging the hole, the male fish began to look around for a mate. After the low tide, the male fish began to dance in front of the female fish to attract the female fish. In order to increase the attraction, male fish often change their bodies from khaki to light taupe, in contrast to black soil. Every male fish tries to introduce the female fish with eggs into his territory and then lure it into his cave. In order to attract the attention of the female fish, the male fish expands his head by expanding his mouth and cheeks, and at the same time, he lures the female fish by bending his back into an arch, erecting his tail fin and constantly twisting his body. If another male fish comes to him, he will try harder to prevent his "the only thrill" from being taken away by others. In the meantime, it will stop every once in a while to see if the other party has lost interest in it or fallen into the clutches of competitors. Then the "suitor" got into the hole and quickly came out to lure the female fish. It seems to convey such a message to the female fish: Come in, this is your warm home. If the female fish still hesitates, she will keep going in and out until the female fish can't resist the temptation and get into its nest. Once the female fish enters its nest, the male fish will return to the hole at a very fast speed and block the "hole" with a piece of mud. Scientists in the spawning room have found that the fertilized eggs of female fish are embedded in the mud wall of the spawning room after fertilization, and it takes about a week for the fertilized eggs to develop on the mud wall. In order to observe the development of eggs and the protective behavior of mudskippers, scientists inserted an endoscopic camera at the top of the spawning room. Through this device, they saw the underground world of mudskippers for the first time. After the fertilized egg matures, the tiny transparent young fish just hatched will attach to the cave water, thus starting their initial trip to sea. In the first few hours after the young fish are born, they look very much like plankton, wrapped in a yellow liquid bag to sustain life. After about 45 days, the young fish can drill out of the cave and come to the beach to start its amphibious life. Towers and trenches We don't know the important role of towers and trenches in biology. We guess that the trench may be used to prevent other animals from approaching their caves, and the tower may be an observation point, from which we can see whether there are prey or carnivores outside the caves. In the morning, we found the mudskipper lazily lying in the tower-shaped hole. At this time, the temperature is 7 degrees Celsius, which is higher than the mudskipper hole. Mud-coated fish are preheated by the hole to prepare for the day's activities. Species classification: mudskipper belongs to Echinocheidae. There are 25 kinds of mudskippers in the world. According to their physical and behavioral characteristics, they can be divided into four species: Scarlaus, Boer Overssalmus, Perez Overssalmus and Perez Overssalmerton. Characteristics of mudskipper There are 3 genera and 6 species in China's coastal areas, namely mudskipper, mudskipper, mudskipper and mudskipper. Common species are mudskipper, big mudskipper and blue mudskipper. Geographical distribution China is mainly distributed in the South China Sea and the East China Sea, and Guangdong is mainly distributed in the Pearl River Estuary and leizhou bay. It is distributed worldwide in the west coast of Africa, Indian, Pacific waters, tropical and subtropical inshore shallow waters of New hebrides (not produced in the United States). Artificial culture 1, pond conditions: generally 2 ~ 10 mu is suitable for breeding mudskippers, and the bottom of the pond should be flat, with soft clay as the best substrate, which is beneficial for mudskippers to burrow and inhabit. The bottom of the pool should be higher than the low tide line in the sea area, which is convenient for drainage and sunning. Dig a cross ditch in the center of the pond and open an annular side ditch along the edge of the pond to form the word "Tian". Generally, the ditch is 2 meters wide and 30 centimeters deep, which is convenient for sheltering mudskippers when draining and drying the lawn. It is best to separate the water inflow and drainage of the pond and connect with the central ditch. The bottom of the pond is inclined from the intake gate to the drainage gate, and the bottom of the drainage gate should be about 20 cm lower than the bottom of the intake gate to facilitate the discharge of pond water. Set up escape prevention facilities around the intake and drainage gates and ponds. 2. Half a month before the swimming pool is cleaned, disinfected and stocked, drain the pool water first and sun the lawn for several days until the lawn surface cracks. Then spray 75 ~ 100 kg of quicklime or 30 ppm of bleaching powder (32% available chlorine) on the whole pond for disinfection to eliminate pests. 3. After the pond for cultivating benthic algae was cleaned and disinfected, we began to cultivate benthic algae. The method is to apply fermented chicken manure or pig manure 20-30 kg, rice bran 15 kg, algae auxin, microorganisms, oil bran, etc. To every acre of pond. When applying fertilizer, choose a sunny day without wind, and scatter chicken manure evenly at the bottom of the pool, being careful not to pile up. Then seawater with the specific gravity of1.010 ~1.01.01.04 will be introduced. After 3 ~ 5 days, benthic algae gradually proliferate and form algae beds. Brown diatoms and green algae are the best algae beds. 4. After sowing and cultivating benthic algae, sowing can be carried out. The seedlings mainly come from natural seedlings collected in the sea area. Choosing fry requires complete fins, scales, no mechanical damage and strong vitality. The seedling density is controlled at 5000 ~ 7000 plants/mu, and the specification is 200 ~ 240 plants/kg. Because the mudskippers don't eat each other, they can be stocked in batches, and the big ones are caught and the small ones are reserved for harvest. 5. Cultivation and management After seeding, the water depth should be kept at about 10 ~ 15cm, and the water quality should be clear, so that sunlight can be transmitted to the bottom of the pond, which is convenient for algae photosynthesis and accelerates the reproduction of benthic algae. However, after a period of cultivation, the original benthic algae will be eaten by mudskippers, and the water quality will become turbid. In order to meet the needs of the growth of mudskippers in the pond, it is necessary to drain water in time to dry the bottom and fertilize it so as to breed benthic algae for the mudskippers to eat. However, it is worth noting that the weather forecast should be paid attention to in the drainage pond, otherwise the algae bed will be washed away in rainy days. In addition, it is also necessary to prevent the harm of fierce fish, birds, crabs and conchs to mudskippers. There are few fish diseases in mudskipper culture, but sometimes parasitic spores appear, with yellow spots under scales and epidermis. Microscopically, it is the cyst of sporozoites, which can be prevented by spraying 0.5ppm "povidone iodine" in the whole pool. There is also a small sea leech, which occasionally parasitizes on thin fish. In severe cases, it can splash 0.5ppm of 90% crystal "trichlorfon" all over the pool. Pest control ① Fish damage: moray eels, sand eels and other fish will prey on mudskippers and block them every time they enter the water to prevent them from invading the pond. ② Bird damage: generally, birds are driven away by driving or warning. (3) Crab damage: Small crabs will invade the mudskipper cave to prey on the mudskipper, or kill the mudskipper by mistake, so the pond should be surrounded by plastic plates to prevent crabs from invading, and small crabs in the pond can also be killed by lights at night. (4) Other hazards: Conch and polychaete will not directly harm the mudskipper, but we should try our best to drive away diatoms in the competition with the mudskipper. You can catch large mudskippers with your hands, cages and nets. The hand-grasping method is that after the fisherman finds the fish hole, he catches the fish directly along the hole with his hands. Its disadvantage is low capture or survival rate. The cage fishing method is that after the fisherman finds the fish hole, he seals the back hole with mud, inserts the fish cage hole into the front hole, and catches it when it comes out of the front hole and enters the fish cage. The net hanging method is to use the habit of large mudskippers in tidal water. First, drain the water in the pond to a depth of 2 cm or dry it, and set a hanging net at the water injection port of the fish pond. When the tide is high, slowly inject water, and the large mudskippers cluster into the hanging net against the current, and pull the net rope to catch them. The main value of mudskipper is small edible fish, with a body length of 50 ~ 90 mm, delicious meat and large output, which has certain economic value. The output of mudskipper is relatively high in China, with an annual output of about 200-300 tons in the coastal beaches of southern Zhejiang, Fujian and Guangdong. Its food chain is short, the breeding cost is low, the investment is low, fish diseases are few, it is easy to breed, and live fish can withstand long-distance transportation. Although the fish is small, its meat is delicious and nutritious, and it has nourishing effect. It is regarded as nourishing food by people in Zhejiang, Fujian, Taiwan Province, Guangdong and other places, and it is in great demand abroad. Mud-coated fish can not only be used as breeding objects in coastal harbor beaches, reclamation areas, mangrove areas and fish ponds, but also be mixed in muddy shrimp ponds. However, in recent 10 years, due to a large number of reclamation, environmental pollution, cold fishing and overfishing in coastal beaches, resources gradually declined, natural seedlings were limited, and the sources were difficult and unstable, which greatly affected the development of aquaculture. In order to protect this resource, especially in the breeding season from May to August, the catch should be limited. In China, the artificial propagation of mudskipper was studied in 1972, 1982, 1986, 1988 ~ 1990, and a certain number of hatched larvae and juveniles were obtained, which provided a scientific basis for future productive seedling raising. In Japan, large-scale mudskippers are distributed in shallow seas and mudflats around Youming Sea in Saga Prefecture, Kyushu, as well as Jianzao Bay and octupole Sea. After seasoning, baking and baking, canned mudskipper is a precious specialty of Saga, so it was selected as the county fish in Saga Prefecture. The annual output of mudskipper in Saga was 2 16 tons in 1964, but it was reduced to 3 tons in 1985 due to overfishing. 1986, in order to save the resources of giant mudskipper, closed fishing periods, closed fishing areas and closed fishing periods were established, and artificial incubation and seedling raising were studied. 1989, Saga successfully released 14000 artificially cultured fry to the mouth of Liujiaochuan River to increase the giant mudskipper resources, which is expected to be recovered soon in Japan. News about Bouncing Fish 20 13 March 14 to 15 Some people in Yingli Town and nearby farms in Leizhou City were poisoned by eating bouncing fish. Wu Feihuai, a poisoned villager in Dingman Village, Yingli Town, Leizhou City, who is being treated at the Second Hospital of Zhanjiang Agricultural Reclamation, said that on the afternoon of March 13, he bought half a catty of fresh jumping fish from Yingli Town Market. After cooking, the couple appeared dizziness, vomiting, numbness of hands and feet and other symptoms soon after eating. When you walk, you stagger like a drunk. "I feel that it may be fish poisoning, and I immediately asked Yingli Town Health Center for help." Wu Feihuai said that in the evening, the doctors in the town health center found that they were deeply poisoned and sent them to the Agricultural Reclamation Hospital overnight. "This kind of jumping fish is highly toxic. I spit the fishbone on the floor. I ate a few chickens at home and they were all poisoned! " Wang Mumei, an employee of Happy Farm, said, I bought jumping fish at Happy Farm Market this afternoon. After eating only a few times, my mouth was closed, my hands and feet were numb and I was unconscious. After my family rushed him to the hospital for emergency treatment, I slowly woke up. Talking about the poisoning situation, Wang Mumei still has a lingering fear. Huang Qingyu, an employee of Happiness Farm, said that all three people in his family were poisoned to varying degrees after eating "jumping fish". His wife and mother are deeply poisoned and are still being treated in the hospital. As of March 7, 20 13, 17, the local hospital * * * received 2 1 person, of which 1 person got worse and was sent to the intensive care unit for treatment, and the other 4 patients recovered and were discharged. On the day of the incident, the main leaders of the city immediately assigned the Municipal Health Bureau, the Ocean and Fisheries Bureau, the Industrial and Commercial Bureau, the Public Security Bureau and the Food Hygiene Administration to quickly intervene in the investigation and do a good job in the treatment of poisoned people. As of March 7, 20 13, 17, the poisoned patient's condition was stable and there was no life-threatening. Later, the provincial CDC judged that the "culprit" of this incident was not "jumping fish" (also known as mudskipper, a kind of goby fish), but goby fish containing tetrodotoxin. Goby goby goby is a kind of poisonous small fish living in warm coastal waters, which contains tetrodotoxin, and fish skin is the most toxic part. In winter and early spring, the toxicity is the highest.
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